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Shannon Diversity Index Calculator

Calculate Shannon-Wiener diversity index, species richness, and evenness from population counts.

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Shannon Diversity Index Calculator

Calculate the Shannon-Wiener diversity index (H), species richness (k), and Pielou evenness (E) from population counts in an ecological community. Enter species names and individual counts to measure how diverse and evenly distributed the community is.

Shannon diversity index formula

The Shannon-Wiener diversity index is defined as:

$$H = -\sum_{i=1}^{k} p_i \ln(p_i)$$

Where $p_i = n_i / N$ is the proportion of individuals belonging to species $i$, $n_i$ is the count for that species, $N$ is the total number of individuals, and $k$ is species richness (the number of species present).

Evenness and related metrics

Pielou's evenness compares observed diversity to the maximum possible for $k$ species:

$$E = \frac{H}{\ln(k)}$$

Evenness ranges from 0 to 1, where values closer to 1 indicate more equal abundance across species. The average population size is simply $N / k$.

Choosing a logarithm base

The natural logarithm (base $e$) is standard in ecology and produces values in nats. Base 2 gives bits (common in information theory), and base 10 gives dits. The relative ranking of communities is unchanged; only the numeric scale differs.

Example calculation

A rainforest survey finds 5 scarlet macaws, 12 blue morpho butterflies, 2 capybaras, 5 three-toed sloths, and 1 jaguar ($N = 25$, $k = 5$):

$$H = -(0.2 \ln 0.2 + 0.48 \ln 0.48 + 0.08 \ln 0.08 + 0.2 \ln 0.2 + 0.04 \ln 0.04) \approx 1.33$$

$$E = \frac{1.33}{\ln 5} \approx 0.83$$

For related ecology tools, see the Carrying Capacity Calculator or the Lotka-Volterra Calculator.

Frequently Asked Questions

What does the Shannon diversity index measure?

It estimates species diversity by combining richness (how many species are present) and evenness (how evenly individuals are distributed among those species). Higher values mean greater diversity.

What is a typical range for the Shannon index?

In real-world ecological data, values usually fall between 1.5 and 3.5 when using the natural logarithm. A value of 0 means only one species is present.

How is evenness different from the Shannon index?

The Shannon index (H) can increase with more species or more equal abundances. Evenness (E) normalizes H by the maximum possible diversity for k species, giving a value between 0 and 1 that reflects only how evenly individuals are spread.

Which logarithm base should I use?

Ecologists typically use the natural logarithm (ln). Base 2 is common in information theory, and base 10 is occasionally used. Choose one base and use it consistently when comparing communities.

What happens if a species has zero individuals?

Species with zero counts are excluded from the calculation. Each row in the calculator should represent a species with a positive population count.

Tags

Shannon Diversity Index Species Richness Evenness Ecology Calculator Biodiversity